Ian Bricknell, Ph.D.

Contact Information

Phone: 207.581.4380

CV: Link to Bricknell CV

Email: ian.bricknell@maine.edu

179 Hitchner Hall
Orono, ME  04469


Ph.D, Lancaster, 1990
BSc, Zoology with Geology, 1st Class Hons, Reading, 1986


Dr. Bricknell comes from North London, in the United Kingdom. Ian initially trained as a medical microbiologist working for the Public Health Laboratory Service, where he received extensive training in human immunology and parasitology. In 1983 he went up to university to read zoology and geology at the University of Reading, under the tutelage of the late Dr L.B. Halstead, graduating with first class honors from there in 1986. During this time he was elected president of the Zoological Society of Reading University and published the first of his 100+ research papers on the diseases of fossil elephants. From Reading University Dr. Bricknell moved to Lancaster University to read for his Ph.D. Here he studied under Prof. W.T.W Potts researching the impact of acid rain on the biology of upland rivers gaining his Ph.D. in 1990. In 1991 he was awarded the Modern Geology Young Geologist of the Year Award for his work on wooly mammoths at the annual British Association for the Advancement of Science meeting.

In 1989 Dr. Bricknell was offered the position of Higher Scientist at Fisheries Research Services (FRS) in Aberdeen in the fish immunology group with Dr. A.E. Ellis as his mentor. Here he worked in the field of fish health developing fish vaccines and new and improved diagnostic methods for the detection of disease in wild and farmed fish. During his first three years at FRS he filed two patents on fish vaccination and improved diagnostic tests on behalf of the Secretary of State for Scotland. In 1993 Dr. Bricknell was promoted to Senior Scientist and in 1999 to Principal Scientist, becoming head of the Immunodiagnostics Department. Ultimately serving on many national and international committees and councils. Including being elected to the Chair of the International Council for the Exploration of the Sea’s Mariculture committee  (term of Office 2005-2008) and received the ICES presidential Service Award in 2008, he has also served on the Norwegian Research Council’s Aquatic Sciences Committee.

In 2007 Dr. Bricknell accepted the post of Libra Professor of Aquaculture Biology at the University of Maine and in 2009 was appointed as the first Director of the Aquaculture Research Institute. Since arriving in the USA he has established a new aquatic animal disease research group. He has expanded his research interests to include lobster health and he is committed to helping the working waterfront in Maine to ensure a sustainable and dynamic aquaculture industry in Maine

Research Interests

Ian Bricknell  joined SMS from his position as Group Leader, Immunological Diagnostics in the Fisheries Research Service Laboratory, Aberdeen, Scotland. He works on the developmental immunity of larval fishes and with host-pathogen interactions. His interests include: the interaction of parasites with their host and the mechanisms they employ to avoid the host’s defense mechanisms; immunological detection of fish diseases; the development of the immune system of larval fishes and the onset of immunocompetence; and, the mechanisms that larval fish use to contain or resist infection.


(Not including symposiums, proceedings etc.)

Michael Pietrak, Alexander Jensen¥, Gayle Barbin Zydlewski & Ian Bricknell. The Three-Spined Stickleback; Gasterosteus aculeatus Linnaeus 1758, Plays a Minor Role as Host of Lepeophtheirus salmonis; (Krᴓyer 1837) in the Gulf of Maine. Journal of Fish Diseases Awaiting DOI

Barker, S.E., Bricknell, I.R., Covello, J., Purcell, S., Fast, M.D., Wolters, W. & Bouchard, D.A. (2019) Sea lice, Lepeophtheirus salmonis (Kroyer 1837), infected Atlantic salmon (Salmo salar L.) are more susceptible to infectious salmon anemia virus. PLOS ONE, 14.

Magnadottir, B., Bragason, B.T., Bricknell, I.R., Bowden, T., Nicholas, A.P., Hristova, M., Gudmundsdottir, S., Dodds, A.W. & Lange, S. (2019) Peptidylarginine deiminase and deiminated proteins are detected throughout early halibut ontogeny – Complement components C3 and C4 are post-translationally deiminated in halibut (Hippoglossus hippoglossus L.). Developmental and Comparative Immunology, 92, 1-19.

Bowden, T.J., Bricknell, I.R. & Preziosi, B.M§. (2018) Comparative pathogenicity of Vibrio spp., Photobacterium damselae ssp damselae and five isolates of Aeromonas salmonicida Var achromogenies in juvenile Atlantic halibut (Hippoglossus hippoglossus). Journal of Fish Diseases, 41, 79-86.

Bricknell, I. (2017) Types of Pathogens in Fish, Waterbourne Diseases. (Chapter 3), In: Fish Diseases Prevention and control strategies (ed. by J. Jeney), pp. 53-74. Wiley, London.

Frederick, C§., Brady, D.C. & Bricknell, I. (2017) Landing strips: Model development for estimating body surface area of farmed Atlantic salmon (Salmo salar). Aquaculture, 473, 299-302.

Piesz, J.§, Barker, S. & Bricknell, I. (2016) Effects of the secretory/excretory products of Lepeophtheirus salmonis on Atlantic salmon leukocyte migration. Fish & Shellfish Immunology, 53, 84.

Borsky, A.J. ¥ & Bricknell, I.R. (2016) The ontogeny of neurosensory structures in larval Atlantic halibut, Hippoglossus hippoglossus (Linnaeus, 1758). Journal of Applied Ichthyology, 32, 669-674

Pert, C.C., Fryer, R.J., Cook, P., Kilburn, R., McBeath, S., McBeath, A., Matejusova, I., Urquhart, K., Weir, S.J., McCarthy, U., Collins, C., Amundrud, T., Bricknell, I.R., 2014. Using sentinel cages to estimate infestation pressure on salmonids from sea lice in Loch Shieldaig, Scotland. Aquac. Environ. Interact. 5, 49-59.


Molloy, S.D., Pietrak, M.R., Bouchard, D.A., Bricknell, I., 2014. The interaction of infectious salmon anaemia virus (ISAV) with the blue mussel, Mytilus edulis. Aquac. Res. 45, 509-518.


Bricknell I, Langston A (2013) Aquaculture: It’s not all about Atlantic Salmon.

J Fisheries Livest Prod 1:e103. doi: 10.4172/2332-2608.1000e103


The settlement and reproductive success of Lepeophtheirus salmonis (Kroyer 1837; Copepoda: Caligidae) on atypical hosts (2012)

Pert Campbell C.; Mordue (Luntz) A. Jennifer; O’Shea Brid; & Bricknell Ian R.

Aquaculture Research Volume: 43   Issue: 6   Pages: 799-805   DOI: 10.1111/j.1365-2109.2011.02891


Bowden, T.J., Bricknell, I.R., 2013. Management of finfish and shellfish larval health in aquaculture hatcheries. Woodhead Publ. Food Sci. Technol. Nutr., 223-245.


Barker, S., Covello, J.M., Bouchard, D., Wolters, W., Fast, M., Bricknell, I., 2013. Lice and ISAV: Are ‘lousy’ salmon more susceptible? Fish Shellfish Immunol. 34, 1637-1638.


Potential role of Mytilus edulis in modulating the infectious pressure of Vibrio anguillarum 02 beta on an integrated multi-trophic aquaculture farm (2012)

Pietrak Michael R.; Molloy Sally D.; Bouchard Deborah A. & Bricknell Ian R.

Aquaculture  Volume: 326   Pages: 36-39   DOI: 10.1016/j.aquaculture.2011.11.024


Molloy SD, Pietrak MR, Bouchard DA & Bricknell IR (2011)

Ingestion of Lepeophtheirus salmonis by the blue mussel Mytilus edulis

Aquaculture   Volume: 311   Issue: 1-4   Pages: 61-64

D. A. Bouchard, D. Basti, and I. Bricknell (2010)

Where Has All The Gaffkemia Gone?

Pages 99-106 In Cipriano, R. C., Bruckner, A., and Shchelkunov, I. S., editors. Aquatic Animal

Health: A Continuing Dialogue Between Russia And The United States. Proceedings

of the Third Bilateral Conference Between the United States and Russia: Aquatic

Animal Health 2009. 12-20 July 2009. Shepherdstown, West Virginia. Michigan

State University, East Lansing, Michigan.

Basti D, Bricknell I, Chang ES & Bouchard D.A. (2010)

Biochemical Reference Intervals For The Resting State In The Adult Lobster Homarus Americanus

Journal of Shellfish Research   Volume: 29   Issue: 4   Pages: 1013-1019

Cook PF, McBeath SJ, Bricknell IR, & Bron JE (2010)

Determining the age of individual Lepeophtheirus salmonis (Kroyer, 1837) copepodids by

measuring stored lipid volume; proof of principle

Journal Of Microscopy Volume: 240   Issue: 1   Pages: 83-86

Basti D, Bricknell IR, Hoyt K & Bouchard DA (2010).

Factors affecting post-capture survivability of lobster Homarus americanus

Diseases Of Aquatic Organisms   Volume: 90   Issue: 2   Pages: 153-166

Kilburn R, Bricknell IR, Cook P, et al (2010).

Design and application of a portable, automated plankton sampler for the capture of the

parasitic copepods Lepeophtheirus salmonis (Kroyer 1837) and Caligus elongatus

(Von Nordmann 1832)

Journal of Plankton Research   Volume: 32   Issue: 6   Pages: 967-970

Urquhart K, Pert CC, Fryer RJ, & Bricknell IR (2010)

A survey of pathogens and metazoan parasites on wild sea trout (Salmo trutta)

in Scottish waters

ICES Journal of Marine Science   Volume: 67   Issue: 3   Pages: 444-453

Patricia Noguera, Catherine Collins, David Bruno, Campbell Pert, Anna Turnbull, Alison McIntosh, Katherine Lester, Ian Bricknell, Stuart Wallace, & Paul Cook (2009)

Red vent syndrome in wild Atlantic salmon Salmo salar in Scotland is associated with Anisakis simplex sensu stricto (Nematoda: Anisakidae)

Diseases of Aquatic Orgnaisms Vol. 87 (3) 199–215

Pert CC, Raffell J, Urquhart K & Bricknell IR (2009)

The pathogen burden of early returning sea trout (Salmo trutta L.) infected with Lepeophtheirus salmonis (Kroyer, 1837), in the River Shieldaig, Scotland

Bulletin of the European Association of Fish Pathologists   Volume: 29   Issue: 6   Pages: 210-216

Gratacap, R.M.L., Thompson, K.D., Bricknell, I.R. & Adams, A. (2009)

Lipopolysaccharide extraction: a phenol alternative.

Journal of Fish Diseases, 32, 811-814.

Basti, D., Bricknell, I., Beane, D. & Bouchard, D. (2009)

Recovery from a near-lethal exposure to ultraviolet-C radiation in a scleractinian coral.

Journal of Invertebrate Pathology, 101, 43-48.

Campbell C. Pert, A. Jennifer Mordue (Luntz), Robert J. Fryer, Bríd O’Shea, Ian R. Bricknell (2009)

The settlement and survival of the salmon louse, Lepeophtheirus salmonis (Krøyer, 1837), on atypical hosts

Aquaculture 288 321–324

McCarthy, U.M., Bron, J.E., Brown, L., Pourahmad, F., Bricknell, I.R., Thompson, K.D., Adams, A., Ellis, A.E. (2008)

Survival and replication of Piscirickettsia salmonis in rainbow trout head kidney macrophages.

Fish & Shellfish Immunology 25, 477-484.

McCarthy UM, Urquhart KL, Bricknell IR (2008)

An improved in situ hybridization method for the detection of fish pathogens

Journal of Fish Diseases 31(9) 669-677

  1. Urquhart, C. C. Pert, R. Kilburn, R. J. Fryer, and I. R. Bricknell (2008)

Prevalence, abundance, and distribution of Lepeophtheirus salmonis (Krøyer, 1837) and Caligus elongatus (Nordmann, 1832) on wild sea trout Salmo trutta L.

ICES Journal of Marine Science, 65, 171-173.


  1. O’Shea, A.J., Mordue (Luntz), R.J Fryer, & I.R. Bricknell 2006

Determination of the Surface Area of a Fish

J Fish Dis 29(7):437-40


I.R. Bricknell, S. Dalesman, B. O’Shea, C.C. Pert, & J. Mordue 2006

The Effect of Environmental Salinity on Sea lice (Lepeophtheirus salmonis) Settlement Success

Dis Aquat Org. 71(3):201-12

McBeath, S. J., Ellis, L. M., Cook, P. F., Wilson, L., Urquhart, K. L. and Bricknell, I. R. 

Rapid development of polyclonal antisera against infectious salmon anaemia virus and its optimisation and application as a diagnostic tool.

J Fish Dis 29 (5), 293–300.

The Sea Bass (Dicentrarchus labrax L.): A Peripatetic Host of Lepeophtheirus salmonis? 2006

C.C. Pert, K. Urquhart and I.R. Bricknell

Bull EAFP 26(4) 163-165

Bergljót Magnadottir, Bjarnheiðu K. Guðmundsdóttir, Sigrún Lange, Slavko Bambir, Agnar Steinarsson, Matthias Oddgeirsson, Tim Bowden, Ian Bricknell, Roy Dalmo & Sigríður Guðmundsdóttir 2006

Immunostimulation of cod (Gadus morhua L.) larvae and juveniles

J Fish Dis 29 147-155


James Bron, Campbell Pert, Christina Sommerville, Hege Helleberg & Ian Bricknell 2006

Diseases of gadoid fish (an update)

Bulletin of the European Association of Fish Pathologists 26(1) 43-46

I.R. Bricknell, J.E. Bron & T.J. Bowden 2005

Diseases of Gadoid Fish in Cultivation: A Review

ICES Journal of Marine Science 63 253-266

Sigrun Lange, Slavko H. Bambir, Alister W. Dodds, Tim Bowden, Ian Bricknell, Sigrun Espelid, Bergljót Magnadóttir 2005

Complement component C3 transcription in Atlantic halibut (Hippoglossus hippoglossus L.) larvae.

Fish & Shellfish Immunology 20 (3) 285-94.

Ian Bricknell & Roy A. Dalmo 2005

The use of immunostimulants in fish larval aquaculture

Fish & Shellfish Immunology 19 (5) 457-472


Bowden, T.J., Butler R. and Bricknell I.R. 2004

Seasonal variation of serum lysozyme levels in Atlantic halibut (Hippoglossus hippoglossus L.)

Fish and Shellfish Immunology Fish & Shellfish Immunology 17(2): 129-135.

David W. Verner – Jeffreys, Robin J. Shields, Ian R. Bricknell and T. Harry Birkbeck 2004

Effects of different water treatment methods and antibiotic addition on larval survival and gut microflora development in Atlantic halibut (Hippoglossus hippoglossus L.) yolk-sac larvae

Aquaculture 232 (1-4): 129-143

I.R. Bricknell, H.E. Hayes, P. Billingsley, A.J. Vigneau, C. Sommerville & R.S. Raynard 2003

Vaccines Against the Salmon lice (Lepeophtheirus salmonis)- A Holy Grail, or just around the corner?

Journal of Parasitology 89(S) 149-157

 Snow, R.S. Raynard, A.G. Murray, D.W. Bruno, J.A. King, R. Grant, I.R. Bricknell, N. Bain & A Gregory 2003

An Evaluation of current Diagnostic tests for the detection of infectious salmon anaemia virus (ISAV) following experimental water-borne infection of Atlantic salmon Salmo salar L.

Journal of Fish Diseases 26, 135-145


I.R. Bricknell, & T.J. Bowden 2003

Immunostimulants and their role in fish health management

Fish Farmer Magazine 26(1) 10-11


Timothy J Bowden, Katy Adamson, Philip MacLachlan, Campbell C Pert and Ian R Bricknell 2003

Long-Term Study of antibody response and injection-site effects of oil adjuvants in Atlantic halibut (Hippoglossus hippoglossus L.)

Fish & Shellfish immunology 14 363-369

David W. Verner – Jeffreys, Robin J. Shields, Ian R. Bricknell and T. Harry Birkbeck 2003

Changes in the gut-associated microflora during the development of Atlantic halibut (Hippoglossus hippoglossus L.) larvae in three British hatcheries.

Aquaculture 219 21-43

T.J. Bowden, D Menoyo-Luque, I.R. Bricknell & H Wegiland 2002

Efficacy of Different Administration Routes for Vaccination Against Vibrio anguillarum in Atlantic Halibut (Hippoglossus hippoglossus L.)

Fish & Shellfish Immunology 12 (3) 283-285


Louisa Wilson, Katy Adamson, Laura Ellis, Paul Cook, Sonia McBeath & Ian Bricknell 2002

An Alkaline Phosphatase-Based Method for the Detection of Infectious Salmon Anaemia Virus (ISAV) in Tissue Culture and Tissue Imprints

Journal of Fish Diseases 25:615-619

I.R. Bricknell, P.F. Bisset & T.J. Bowden 2002

The Function of Atlantic Salmon (Salmo salar L.) Antibodies Under Extremes of pH and Osmolarity.

Fish & Shellfish Immunology 13:215-219

Hiney M., Rodger H., Bricknell I., Casburn P. & Mulcahy D. 2002

Aquaculture development and regulation: incompatibility or harmony?

Bulletin of the European Association of Fish Pathologists 22:178-184

Robert S Raynard, Ian R Bricknell, Peter F Billingsley, Alasdair J Nisbet, Antoine Vigneau and Christina Sommerville 2002

Development of vaccines against sea lice

Pest Management Science 58:569-575

D.A. Smail, R. Grant, K. Ross, I.R. Bricknell & T.S. Hastings 2001

The use of haemabsorption for the isolation of infections salmon anaemia virus on SHK-1 cells from Atlantic salmon (Salmo salar L.) From Scotland

Bulletin of the European Association of fish Pathologists 20(5) 212-214


  1. J. Bowden, K. Lester, P. MacLachlan and I. R. Bricknell (2000)
    Preliminary Studies into the Short-Term Effects of Adjuvants on Atlantic Halibut (Hippoglossus hippoglossus L.)

Bulletin of the European Association of fish Pathologists 20(4) 148-152

  1. Snow, C. O. Cunningham I. R. Bricknell. (2000)

Susceptibility of Juvenile Atlantic Cod (Gadus morhua) to Viral Haemorrhagic Septicaemia Virus Isolated from Wild Caught Atlantic Cod.

Diseases of Aquatic Organisms 41:225-229

 Bricknell I.R., Bowden, T.J., Verner-Jefferys D.W., Bruno D.W., Shields R.S., and Ellis A.E. (2000)

Susceptibility of Juvenile and sub-adult Atlantic halibut (Hippoglossus hippoglossus, L.) to infection by Vibrio anguillarum and efficacy of protection induced by vaccination.

Fish & Shellfish Immunology 10:319-327


Ron Stagg, David Bruno, Carey Cunningham, Trevor Hastings & Ian Bricknell (1999)

Infectious Salmon Anaemia: Epizootiology and Pathology.

State Veterinary Journal 9(3) 1-5

Trevor Hastings, Gilles Olivier, Roland Cusack, Ian Bricknell, Åre Nylund, Martin Binde, Pauline Munro, Charles Allen (1999)

Infectious Salmon Anaemia

Bulletin of the European Association of fish Pathologists 19(6) 286-288

Bricknell I.R. Bowden, T.J., Bruno D.W., MacLachlan P., Johnstone R. and Ellis A.E. (1999)

Susceptibility of Atlantic halibut, Hippoglossus hippoglossus, (L.) to infection with typical and atypical Aeromonas salmonicida.

Aquaculture 175 1-13


Bricknell I.R. Bowden, T.J., King J. A. & Ellis A.E. (1999)

Duration of Protective Antibodies and the Correlation with Protection in Atlantic Salmon (Salmo Salar L.) Following Vaccination with An Aeromonas Salmonicida Vaccine Containing Iron Regulated Outer Membrane Proteins and Secretory Polysaccharide.

Fish Shellfish Immunol. 9 (2)139-151

O’Dowd A.M., Bricknell I.R., Secombes C.J., & Ellis A.E.(1999)

The Primary and Secondary Antibody Responses to Iron Restricted Outer Membrane Proteins Antigens Atlantic Salmon (Salmo salar L.) Immunised with A-layer Positive and A-layer Negative Aeromonas Salmonicida Bacterins)

Fish Shellfish Immunol. 9 (2) 125-138

Ferguson Y., Bricknell I.R., Glover L.A., MacGregor D.M., and Prosser J.I. (1998)

Transmission of lux-Marked and wild type Aeromonas salmonicida strains between Atlantic salmon (Salmo salar L.)

FEMS Microbiology Ecology vol. 27 pp. 251-260


  1. Vipond, I.R. Bricknell, Emma Durant, T.J. Bowden, A.E. Ellis, Mary Smith and S.M. MacIntyre, (1998)

Defined deletion mutants can identify the major virulence factors of Aeromonas salmonicida.

Infection and Immunity, May 1998, p1990-1998


I.R. Bricknell, T.J. Bowden and A.L. Langston, (1997)

Halibut Health and Immunology

Bulletin of the European Association of Fish Pathologists 17(6) 267-269


Bowden, T.J., Butler R., Bricknell I.R. and Ellis A.E. (1997)

Serum trypsin-inhibitory activity in five species of farmed fish

Fish and Shellfish Immunology 7, 377-385


Smail D.A. Bricknell I.R. and Patterson D. (1997)

The inability to detect Infectious Pancreatic Necrosis Virus (IPNV) in crustacean diets for  larval Atlantic halibut, Hippoglossus hippoglossus L..

Bulletin of the European Association of Fish Pathologists 17(4) 129-131

Bricknell I.R. Bowden T.J. Lomax, L. and Ellis A.E. (1997)

Antibody response and protection of Atlantic salmon (Salmo salar) immunised with an extracellular polysaccharide of Aeromonas salmonicida

Fish and Shellfish Immunology 7 (1) 1-16

Bricknell I.R., Bruno D. W. and Stone J. 1996

Aeromonas salmonicida infectivity studies in goldsinny wrasse Ctenolabrus rupestria, L.

Journal of Fish Diseases 19, 469-474

  1. R. Bricknell, D. W. Bruno, T.J. Bowden and P. Smith 1996

Fat Cell Necrosis Syndrome in Atlantic Halibut (Hippoglossus hippoglossus L.).

Aquaculture 144 65-69

Bricknell I.R. and Bruno D. W. 1995

Sudden fish death associated with Alcian Blue marking.

Bulletin of the European Association of Fish Pathologists 15(3) 1‑2

Bricknell I.R. 1995

A reliable method for the induction of experimental furunculosis

Journal of Fish Diseases 18(2) 127‑133

R.S. Raynard, A.L.S. Munro, J. King, A. E. Ellis, D. W. Bruno, I. R. Bricknell, P. Vahanakki, R. Wotton, C. Sommerville, A. Petrie, B. Vivers, O. Andrade‑Salas, W. Melvin, T. Amezega, M. B. Labus, J.J. Coull, P. Relly, M.F. Mulcahy, M. O’Donoghue, and J. O’Connell. 1994

Development of a vaccine for the control of sea lice (Lepeophtheirus salmonis and Caligus elongatus) in Atlantic salmon (Salmo salar)

ICES CM 1994/F:7

Bricknell I.R. & Ellis A.E.  1993

The development of an effective vaccine against furunculosis

Fish Farmer 16(5) 16‑19


Coleman G., Bennett A.J., Whitby P.W. and Bricknell I.R.   1993.

A 70 kDa Aeromonas salmonicida serine protease ‑ b‑galactosidase hybrid protein as an antigen and its protective effect on Atlantic salmon (Salmo salar L.) against a virulent A. salmonicida challenge.

Biochemical Society Transactions 21(49S)

Ellis A.E., Bricknell I.R. & Munro A.L.S. 1992.

Patent PCT/GB92/01016 Vaccines against Aeromonas salmonicida

Bricknell I.R., Ellis A.E. & Munro A.L.S. 1992.

Patent D/IPR/1G0024 Methods for the detection of the protective antigens of Aeromonas salmonicida.

Bricknell I.R., Ellis A.E. & Munro A.L.S. 1992.

The Current Status of a Scottish Furunculosis Vaccine.

ICES CM 1992/F:29

Ellis A.E., Bricknell I.R. & Munro A.L.S. 1992.

The Status of Furunculosis Vaccines.

Bulletin of the Aquaculture Association of Canadian 92(1) 10‑14

Potts W.T.W. and Bricknell I.R. 1992.

The Ionic Composition of Ion Transporting Cells in Stonefly Nymphs.

Scanning Microscopy 6(1) 287‑294

Bricknell I.R. 1991.

X‑Ray Microanalysis as a Tool for Measuring Intracellular Ions

Microscopy and Analysis 5(3) 7‑13

 Bricknell I.R. & Potts W.T.W., 1989

Silver Staining of Ion Transporting Cells

Practical Exercises in Biology Vol. 7 (W.T.W. Potts Ed.) Lancaster University Press, Lancaster.

Bricknell I.R. & Potts W.T.W., 1989

The Distribution of the Mitochondria Rich Ion Transporting cells of British Plecoptera, In relation to the Habitat

Entomologist 108(3) 176‑183

Bricknell I.R. & Potts W.T.W., 1988

The Effects of Acidification on the Coniform Mitochondria Rich Ion Transporting Cells of Plecoptera

Proceedings of the Royal Microscopic Society 23 107‑108

Bricknell I.R. & Potts W.T.W. 1988

The Effects of Low pH on the Coniform Mitochondria Rich Ion Transporting Cells of Plecoptera.

Institute of Physics Conference Series 93(3) 583‑584

Bricknell I.R. 1986.

The Paeleopathology of Pleistocene Proboscideans

Modern Geology 11(4) 186‑202              

Invited papers/plenary lectures

Sea Lice Secreted Products Can Reduce Lepeophtheirus salmonis Infection in Atlantic Salmon
Jessica L. Piesz, Sarah E. Barker, Ian R. Bricknell WAS New Orleans March 2019

I Bricknell. Disease Ecology in Integrated aquaculture systems Second international symposium on IMTA Busan University Republic of Korea 14-15th September 2018

Ian Bricknell. A Physiological Basic for sea lice vaccines. John Hopkins University Invited Lecture Series May 2018

Ian Bricknell. Why are sea lice such a successful parasite in farmed fish? the Institute of Marine Environmental Technology Public Lecture Series Baltimore May 2018

Ian Bricknell. 30 Years in the trenches. Where did we go wrong (and Right) in Sea Lice research? 44th  Eastern Fish Health Workshop Chattanooga TN 9-14th April 2018

Ian Bricknell and Jessic Piezs “Why Current Vaccines Against Sea Lice Don’t Work” Aquaculture America San Antonio Texas Feb 2017

Ian Bricknell “Hunting the Bugs, disease ecology in a simple ecosystem” Inchon Technical University South Korea July 2017

Ian Bricknell “Aquaculture in a changing Environment” 1st International symposium on Integrated Multi Trophic Aquaculture, Busan South Korea August 2017

I Bricknell. That’s Because you are an Evil Aquaculture Scientist. World Aquaculture 2016 Las Vegas Nevada Feb 22-26  2017

Ian Bricknell “Advancing knowledge in IMTA via information exchange and cooperative research between USA and Korea.” Annual JPA meeting University of Washington January 2016

I Bricknell & Emma Taccardi “The Lumpfish Cyclopterus lumpus as a Potential Biological Control for Sea Lice on Salmon Farms” 40th Eastern Fish Health Conference, Virginia Sept 2015

Where are all the sea lice? Searching the wild fish of Cobscook Bay

  1. Jensen, M. Pietrak*, S. Barker, G. Zydlewski and I. Bricknell

Sea Lice 2014 Aug31st-Sept 5th Portland Maine

  1. Frederick*, M. Pietrak, S. Barker, D. Brady and I. Bricknell

Where are all the sea lice? A first glance at sentinel fish in Cobscook Bay

Sea Lice 2014 Aug31st-Sept 5th Portland Maine

J.L. Piesz*, I.R. Bricknell, Hernan Pizarro, S.E. Barker

Increased susceptibility to infectious salmon anemia virus (ISAv) in Lepeophtheirus salmonis – infected Atlantic salmon

Sea Lice 2014 Aug31st-Sept 5th Portland Maine

A physiological approach to vaccines against fish parasites

Ian Bricknell1, Sarah Barker1 and James Bron2

1 Aquaculture Research Institute 2 5735 Hitchner hall, University of Maine Orono, ME 04469-5735, 2 Institute of Aquaculture University of Stirling, Stirling, UK FK9 4LA


Disease risks on integrated multi-trophic aquaculture farms: reshaping disease paradigms Michael Pietrak*, Sally Molloy, Deborah Bouchard, and Ian Bricknell Aquaculture Research Institute University of Maine 5735 Hitchner Hall Orono, ME 04469-5735



Ecological aquaculture in land-based and open water systems or How to Manage
Diseases in Artificial Ecosystems I Bricknell Bremerhaven Forum February 2013

Barker, S., Covello, J.M., Bouchard, D., Wolters, W., Fast, M. and Bricknell, I.

Sea Lice and ISAv: Are ‘lousy’ salmon more susceptible?

1st International Conference of Fish and Shellfish Immunology, Vigo, Spain, June 25- 28, 2013.

Barker, S., Bouchard, D., Covello, J.M., Wolters, W., Fast, M. and Bricknell, I.

Are ‘lousy’ fish more susceptible to ISAV infection? And what is the role of lice in

transmitting ISAv?

Atlantic Canada Fish Farmers Association Annual Fall Workshop, St Andrews, N.B.,

November 14-15, 2012.

Covello, J.M., Barker, S., Bouchard, D., Wolters, W., Fast, M. and Bricknell, I.

Effects of Sea Lice Infections and ISAv on Atlantic Salmon: host strain differences.

Atlantic Canada Fish Farmers Association Annual Fall Workshop,

St Andrews, N.B., November 14-15, 2012.

T.J. Bowden and I.R. Bricknell. Getting to the point. How to develop an injectable vaccine.

Eastern Fish Health Workshop, Gettysburg. April 2013

I Bricknell & M Pietrak Gangneug Sea Grant International Symposium”

Held at Gangneug University Republic of Korea March 26th 2013

Vaccines against fish parasites

I Bricknell” 1st International Integrated Multi-Trophic Aquaculture (IMTA) Symposium”

Busan, Republic of Korea March 28-29, 2013

Pietrak M, Bouchard, D. & Bricknell I

Preparation of finfish for disease diagnostics


Finfish Culture in Recirculating Aquaculture Systems: Let’s Talk Integration

M Pietrak & I Bricknell

NACE 2013 Mystic Connecticut



Disease Risks on Integrated Multi-Trophic Aquaculture Farms: Reshaping Disease Paradigms

M Pietrak D Bouchard Sally Dixon-Molloy & I Bricknell


NACE 2013 Mystic Connecticut

Deploying a Commercial Scale Mussel Raft on a Salmon Farm: More than just growing fish and mussels

Mike Pietrak, Chris Bartlett, Dana Morse, Jeff Robinson, Sally Molloy, Deborah Bouchard, David Miller, and I Bricknell.


NACE 2013 Mystic Connecticut

Gulf of Maine Research Institute SeaState Lecture 7.0 Series March 8, 2012

Integrated Multitrophic Aquaculture. How to Manage Diseases in an Artificial Ecosystem

Ian Bricknell, (invited speaker)

Scratch My Itch: Those Pesky Sea Lice

Dr Ian Bricknell (invited speaker)

37th Eastern Fish Health Workshop; High Peaks Resort, Lake Placid, New York 23 – 27 April 2012


 2011 First EWOS Fish Biotechnology Lecture, Immunostimulants a new dawn in fish vaccines?

Ian Bricknell

PEI Veterinary School May 2011

2010 Plenary Lecture Immunology of Fish Parasites,

Ian Bricknell

1st EOFFI Conference Viterbo Italy June 3-8

2009 Plenary Lecture Prevalence of Gaffkemia in Pounded Lobsters. Third Bilateral Conference Between the 2009 David Basti, Deborah Bouchard & Ian Bricknell

Distribution of Sea Lice in Enclosed Bays, PEI Veterinary School Lecture Series

United States and Russia: Aquatic Animal Health 2009 July 12 – 20, 2009 Clarion Hotel and Conference Center Shepherdstown, West Virginia, U.S.A.

2008 GMRI Sea State 3.0 – Sea Lice: Do They Come From Fish Farms?

Ian Bricknell

March 15th GMRI Portland ME


Keynote lecture: Monitoring the Infective Pressure of Lepeophtheirus Salmonis (Krøyer 1837) on Wild Salmonid Populations in Loch Torridon, Scotland

Ian R. Bricknell, Campbell C. Pert, Paul Cook, Rachael Kilburn, Sonia McBeath

World Aquaculture Society San Antonio Texas Feb 27-March 3rd 2007 Special session SEA LICE- Infection Dynamics on Farms and Interactions with Wild Salmon

Plenary lecture: The physiology of sea lice (L. salmonis) in relation to successful settlement on its host’s

I.R. Bricknell & J.E. Bron

ICOPA XI, The Scottish Exhibition and Conference Centre (SECC), Glasgow Scotland UK.

6-11th August 2006

Sea Lice infection of salmonid smolts in estuaries

I.R. Bricknell

Invited speaker: University of Aberdeen Seminar series May 2006

Flexibacter ovolyticus and other emerging pathogens in the UK halibut industry

I.R. Bricknell

Plenary lecture BMFA VIIIth annual conference, November 2006 Date & location TBC)

Biosecurity in ornamental aquaculture: keeping Koi Herpes virus out

I.R. Bricknell

Plenary lecture: OATA conference,13th-14th November 2006 The Britannic Royal Hotel, Coventry

Selected recent papers/plenary lectures

Advances in Vibrio anguillarum 02b vaccines in farmed Atlantic cod

I.R. Bricknell & K. Urquhart

Plenary lecture BMFA annual conference, Oban, November 8-9 2005

Rapid Detection of BKD in Salmonid Fish by IFAT

  1. Urquhart٭, P. Cook, S. J. McBeath, J. E. Bron, and I. R. Bricknell

11th International meeting of the EAFP Copenhagen September 2005

Workshop 4 Diseases of gadoid fish

Ian Bricknell & James Bron (coordinators)

11th International meeting of the EAFP Copenhagen September 2005

0-119 Emergence of Disease risks to gadoid in culture

Ian Bricknell, Tim Bowden, S.C. Johnson & James Bron

11th International meeting of the EAFP Copenhagen September 2005

0-120 Parasites of Cultured gadoids: current perspectives and future prospects

James Bron, A.P. Shinn, S.C. Johnson & Ian Bricknell

11th International meeting of the EAFP Copenhagen September 2005

The Survival of the parasitic sea louse (Lepeophtheirus salmonis) to the infectious copepodid stage and their survival on atypical hosts

Campbell Pert, A. Jennifer Mordue (Luntz) & Ian Bricknell

11th International meeting of the EAFP Copenhagen September 2005

Immunostimulants in Aquaculture

Ian Bricknell, Tim Bowden  & Roy Dalmo

Plenary lecture Aquatech Chile, Puerto Monte August 2005

Sea Lice (Lepeophtheirus salmonis) A European problem?

Ian Bricknell & James Bron

Plenary lecture Aquatech Chile, Puerto Monte August 2005

Dealing with Emerging Fish Disease: Gadoid Diseases

Ian Bricknell, James Bron, Katy Urquhart & Tim Bowden

Plenary lecture Aquatech Chile, Puerto Monte August 2005

Post Settlement Dynamics of the Parasitic Sea Louse, Lepeophtheirus salmonis (Copepoda: Caligidae)

Brid O’Shea, A. Jennifer Mordue (Luntz) & Ian Bricknell

9th international conference on copepoda, Tunis, Tunisia, July 2005

Sea Lice (Lepeophtheirus salmonis) Settlement Success at Low Salinity

Ian Bricknell, Sarah Dalesman, Brid O’Shea, Campbell Pert & A. Jennifer Mordue (Luntz)

9th international conference on copepoda, Tunis, Tunisia, July 2005

Growing Sea Lice in the laboratory

Ian Bricknell & James Bron

Plenary Lecture

9th international conference on copepoda, Tunis, Tunisia, July 2005

The Use of Genomics, Proteomics and Immunology in Studies of the Parasitic Copepod (Lepeophtheirus salmonis)

Fast MD, Bron JE, Bricknell IR, Ross NW, Johnson SC

Plenary Lecture

9th international conference on copepoda, Tunis, Tunisia, July 2005

I.R. Bricknell

Disease of Gadoid fish: A Review

Plenary Lecture

BMFA VIIth Annual Scientific Meeting

Oban 2-3 November 2004

Disease Of Gadoid Fish In Cultivation: What Do We Know And What Do We Need To Know?

I.R. Bricknell & T.J. Bowden

Plenary Lecture

ICES Mariculture of Gadoids Conference

Bergen June 2004

Salmon Lice Settlement Success at Low Salinity

I.R. Bricknell, S. Dalesman & J.A. Mordue

6th Sea Lice Conference Huntsman Marine Science Centre St. Andrews New Brunswick Canada June 2003

Settlement of Salmon Lice on “Dead End” Hosts

I.R. Bricknell, C.C. Pert & J.A. Mordue

6th Sea Lice Conference Huntsman Marine Science Centre St. Andrews New Brunswick Canada June 2003

The physiological environment found in the gut of the sea louse (L. salmonis)

I.R. Bricknell, J.E. Bron, P. Cook, K. Adamson, A.J. Vigneau, R.S. Raynard and P.F. Billingsley

10th EAFP Meeting Malta September 2003

Winner of a 2003 EAFP Best Poster Award

Do Infected Salmon Recruit More Sea Lice, Lepeophtheirus salmonis, Than Naïve Fish?

Bríd O’Shea, A.J. Mordue (Luntz) I.R. Bricknell

10th EAFP Meeting Malta September 2003

The Fecundity Of The Salmon Louse, Lepeophtheirus salmonis and Their Survival On Atypical Hosts

Campbell C. Pert, A. Jennifer Mordue (Luntz) and Ian R. Bricknell

10th EAFP Meeting Malta September 2003

Host Preferences of the Salmon Louse, Lepeophtheirus salmonis

Bríd O’Shea, A.J. Mordue (Luntz), I.R. Bricknell

6th Sea Lice Conference Huntsman Marine Science Centre St. Andrews New Brunswick Canada June 2003

The Impact of Salinity on Survival of Lepeophtheirus salmonis

S.Dalesman, I.R. Bricknell, C. Pert* B. O’Shea and A.J. Mordue (Luntz)

6th Sea Lice Conference Huntsman Marine Science Centre St. Andrews New Brunswick Canada June 2003

 Ready Salted: Salmon Louse Aminopeptidase

A.J. Vigneau, I.R. Bricknell, R.S. Raynard and P.F. Billingsley

6th Sea Lice Conference Huntsman Marine Science Centre St. Andrews New Brunswick Canada June 2003

Vaccines Against Sea Lice: A Holy Grail or Just Around the Corner?

I.R. Bricknell

Plenary Lecture

ICOPA X Canada House, Vancouver, Canada July 2002