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JoyceJoyce E. Longcore. Joyce received a B.S. in Biology from the University of Michigan in 1960 and worked during the following year for Professor F.K. Sparrow, eminent student of the zoosporic fungi. She then attended Indiana University under the direction of Sparrow’s former Ph.D. student, Robert Johns. Longcore received a Masters of Arts and returned to Ann Arbor to work for Professor Sparrow before marrying in 1964. After marriage, she worked for Sparrow from the University of Maryland laboratory of another of his former students, Robert Paterson. She then became a full-time housewife and mother until beginning graduate school at the University of Maine in 1984. Dr. Richard Homola, student of Alexander Smith and a specialist of Basidiomycetes, was her major professor while she pursued a Ph.D. studying chytrids. She received a doctorate in 1991 and was fortunate in being allowed to maintain space in the then Department of Botany and Plant Pathology. Currently she is an Research Scientist in the School of Biology & Ecology, where she studies zoosporic fungi and maintains cultures of diverse chytrids and their allies plus cryopreserved isolates of the amphibian pathogen Batrachochytrium dendrobatidis. She furnishes cultures for research and teaching.


Former students

jeffp-cropJeffrey S. Piotrowski. Jeff graduated from the University of Georgia and came to the University of Maine, where Seanna Annis and Joyce Longcore advised him for his masters work. He received his degree in 2002 with a thesis entitled “Physiology, enzyme production, and zoospore behavior of Batrachochytrium dendrobaditis, a chytrid pathogenic to amphibians.” Jeff is now a scientist at the DOE Great Lakes Bioenergy Research Center.



Marilyn R. N. MolliconeMarilyn R. N. Mollicone, author of Monoblepharidales pages at this site. Marilyn Rose Noyes grew up on a farm in Bethel, Maine and learned the beginnings of natural history from her mother and father. While still in grade school, a program of color slides of native flowers by a local woman photographer inspired her lifelong passion for botany. She attended Gould Academy in Bethel, Maine and graduated in 1946, second in her class. Marilyn attended the University of Maine where she was the only woman in most of her classes in the College of Agriculture. She graduated in 1950 with a B.S. in Botany, with High Distinction. Later in 1950, Marilyn married Phillip Mollicone and soon gave up her studies to start a family. While her son and daughter were teenagers, she began a career as a naturalist as the director of a natural history day camp program for children held at the Augusta Nature Center, Augusta, Maine, where she remained the director for 19 years. Marilyn’s husband, Phillip, died in 1978. Eight years later, in 1986, at the age of 58, Marilyn returned to the University of Maine where her mentor was Dr. Joyce Longcore, student of chytrids. She chose to study the Order Monoblepharidales when she found a member of the order in her first sample from nature, collected among cattails growing in a shallow pond in an old quarry in Augusta Nature Center. Marilyn received her Master of Science in Botany and Plant Pathology in 1993. Her thesis was entitled “Life Cycle, Morphology, Culture, and Zoosporic Ultrastructure of Monoblepharis polymorpha”. Her initial Ph.D. research focused on the ultrastructure of Gonapodya polymorpha, which was published as “Zoospore ultrastructure of Gonapodya polymorpha” in the Journal Mycologia in 1999. She isolated and grew pure cultures of several genera of Monoblepharidales, studied details of their development and photographed many specimens of the order. Unable to continue her Ph.D. work because of health issues, she concentrated on preparing directions to enable others to obtain and use members of the Monoblepharidales for teaching and research.

Rabern-croppedD. Rabern Simmons. Rabern graduated from the University of Virginia’s College at Wise. He began a master’s degree program at the University of Maine in January of 2005 and received his degree in May of 2007 with a thesis entitled “Systematics of the Lobulomycetales, a new order within the Chytridiomycota.” Rabern continued at UMaine in a Ph.D. program supported by the NSF PEET program. He received his PhD in 2012 with a dissertation entitled “Systematics of the Powellomycetaceae Fam. Nov and the detection of Spizellomycetalean chytrids from manure and soil substrates.” After a post doc working with Hirsutella, a genus that contains a fungal pathogen of ants with Dr. Ellie Groden at the University of Maine , he is now working as a post doc on fungi associated with ambrosia beetles in the lab of Jiri Hulcr at the University of Florida.


Sasha E. Greenspan. Sasha graduated from Wesleyan University. At the University of Maine’s program in Ecology and Environmental Sciences she was co-advised by Joyce Longcore (School of Biology & Ecology) and Aram Calhoun (Department of Wildlife Ecology). She received a masters degree in 2011 with a thesis entitled “Establishment of Batrachochytrium dendrobatidis in anuran epidermis and experimental transmission from bullfrogs to wood frogs.” She is now in a PhD program with Ross Alford at James Cook University.